A new subfamily of linyphiid spiders based on a new genus created for the <i>keyserlingi-group </i>of the genus <i>Lepthyphantes</i> (Aranei: Linyphiidae)
A new linyphiid subfamily, Ipainae subfam.n., is established to include seven genera and 26 species, of which one genus is new to science: (1) The type genus <i>Ipa </i>gen.n., containing <i>I. keyserlingi </i>(Ausserer, 1867), comb.n. (the type species), <i>I</i>. <i>terrenus </i>(L. Koch, 1879), comb.n. -<i> I. quadrimaculatus </i>(Kulczynski, 1898), syn.n., comb.n., <i>I</i>. <i>spasskyi </i>(Tanasevitch, 1986), comb.n. and. <i>pepticus </i>(Tanasevitch, 1988), comb.n., all ex <i>Lepthypahntes; </i>(2) <i>Epibellowia </i>Tanasevitch, 1996, containing <i>E. septentrionalis </i>(Oi, 1960) (the type species), <i>E. pacificus </i>(Eskov & Marasik, 1992) and <i>E. enormitus </i>(Tanasevitch, 1988); (3) <i>Metalepthyphantes </i>Locket, 1968 <i>ad partem, </i>containing <i>M</i>. <i>bifoliatus </i>Locket, 1968, <i>M. carinatus Locket, </i>1968, <i>M. clavator </i>Locket, 1968, <i>M. dentiferens </i>Bosmans, 1979, <i>M. machadoi </i>Locket, 1968 (the type species), <i>M. ovatus </i>Scharff, 1990, <i>M. perexiguus </i>(Simon & Fage, 1922), <i>M. praecipuus </i>Locket, 1968 and <i>M. vicinus </i>Locket, 1968; (4) <i>Solenysa </i>Simon, 1894, containing <i>S. circularis </i>Gao, Zhu & Sha, 1993, <i>S. geumoensis </i>Seo, 1996, <i>S. longqiensis </i>Li & Song, 1992, <i>S. melloteei </i>Simon, 1894 (the type species), <i>S. protrudens </i>Gao, Zhu & Sha, 1993 and <i>S. wulingensis </i>Li & Song, 1992; (5) <i>Uralophantes </i>Esynin, 1993, containing <i>U. troitskensis </i>Esynin, 1993 (the type species); (6) <i>Wubanoides </i>Eskov, 1986, containing <i>W. uralensis </i>(Pakhorukov, 1981) -<i> W. longicornis </i>Eskov, 1986 (the type species) and <i>W. fissus </i>(Kulczynski, 1926); and (7) <i>Epigytholus </i>Tanasevitch, 1996, containing <i>E. tuvensis </i>Tanasevitch, 1996 (the type species). Because both Micronetinae and some representatives of the new subfamily Ipainae have the socalled Fickert's gland inside their radix, they are hypothesized to be sister groups despite otherwise greatly deviating secondary genital organs.