A New Primitive Therocephalian (Therocephalia, Theromorpha)from the Middle Permian of Eastern Europe
The largest scylacosaurid therocephalian Julognathus crudelis gen. et sp. nov. is described based on materials from the middle Permian locality of Sundyr-1, Republic of Mari El, Russia. In Sundyr tetrapod community, Julognathus occupied the niche of the largest predator, forming a trophic pair with a herbivorous tapinocephalian.