THE ORIGIN OF VESSELS AND VASCULAR PLANTS
Original and literature data about the structural aspects of symbiogenetic acts which preceded to appearance of vascular plants and continent colonization by them and the conception based on them are described. Two symbiogenetic acts (prochlorophytes + protists - marine algae; marine algae + fungi - land vascular plants) are presented in plant body by two water transport networks originated as the buffer zones of symbiotic exchange. Descending phloem arises in phylogenesis from the membrane capsula of prochlorophytes. Rising xylem is the special part of the apoplast. It starts from mycelium channels of fungi. Conducting elements of phloem and xylem are formed by transport fluxes which are not gomologous: cyanobacteria photosynthesis is the source of phloem flux, fungal proteolysis is the same for xylem flux. It is shown that the disintegration of symbiotic partnership in one of pares is accompanied by the reduction of transport network corresponding to it. Basal steps of vascular plant evolution are reflected by the structure of conducting tissues and transport fluxes. Anatomical reconstruction of the origin and history of vascular plant migration is given on this theoretical base.